Darwins Argument
individuals whose inherited traits give them a higher probability of surviving and reproducing in a given environment tend to leave more offspring than other individuals
--> leads to the accumulation of favorable traits in the pop. over generations
Natural Selection
Interactions between the environment and organismal variation such that individuals with certain inheritable characteristics survive and reproduce at a higher rate than other individuals
Common Descent
One origin of life, or multiple origins.
Homology
Similarity due to common origin (Darwin's belief)
Ontogeny
History of the development of an organism through its entire life. (from birth --> child --> adult)
Recapitulation
Repeating the evolutionary stages of its ancestors during its own embryonic development. (Ontogeny recapitulates phylogeny)
Heterochrony
Evolutionary change in timing of development
- shortening or lengthening of ontogeny results in a mosaic of different kinds of developmental evolutionary change in a single lineage
Allopatric Speciation
Occurs when populations occupy separate geographical areas. (Physical Barrier, become 2 species)
Type of Reproductive Barrier
Sympatric Speciation
Process of a new species evolving from a single ancestral species while living in the same environment but faced with different components.
Parapatric Speciation
Geographically intermediate between allopatric and sympatric speciation. (Same spot just in different locations, have a limited border)
Gradualism
Evolution occurs by the accumulation of small changes over time.
Punctuated Equilibrium
May explain the discontinuous evolutionary changes observed through geological time.
- Gradual change then sudden change over thousands of years
Exaptation
Evolutionary cooption of a biological character for a role unrelated to the character's evolutionary origins
- structures are used for something else, than initially intended for
Neo-Darwinism
Combines knowledge about genes, chromosomes and inheritance with Darwin's theory
Population Genetics
Studies evolution as change in gene frequencies in populations. (looking at genes now --> overtime)
Microevolution
Evolutionary changes in frequencies of different allelic forms of genes.
Macroevolution
Evolution on a grand scale with origins of new structures and designs, trends, mass extinctions, etc.
Evolution
Changes in allele frequencies (measurable --> can see overtime)
Mutation
Change in DNA sequence (due to error in DNA replication and external sources like chemicals, radiation, or U.V. sunlight)
Gene Flow
Transfer of alleles between populations (genes moving together, can see the diff in genes and alleles from parents)
Genetic Drift
Occurs when chance events alter allele frequencies (typically happens with a small populations) --> based on luck not fitness
Founder Effect
A loss of genetic variation that happens when a new population is started by a small group of individuals from a larger parent population. (ex. amish)
Bottleneck Effect
A sharp reduction in population size and genetic diversity caused by a random event. (survival is based on luck not traits)
Intrasexual Selection
Selection within the same sex.
Intersexual Selection
(mate choice) Individuals of one sex are choosy in selecting their mate from the other sex.
Colonial Organization
An aggregation of undifferentiated cells. Cells all have the same function.
Multicellular Organization
Aggregation of cells that are functionally differentiated. Division of labor, each cell has a different function: reproduction, digestion, respiratory, etc.
What symmetry do humans classify as?
Bilateral
Radically Symmetrical
An arrangement of body parts or shapes around a central axis. Object can be divided into identical halves.
Asymmetrical
Having 2 sides or halves that do not match or are not the same size, shape, and arrangement.
Cephalization
Sensory apparatus, concentration of sensory organs at the front or anterior end (to form a head).
Ex. jelly fish have organs everywhere --> no sense of direction, humans have organs facing front (associated with bilateral animals--> move in one direction)
Cleavage
Initial cell division of zygote
Body Cavities
Space for organs, fluids, etc
Increases SA and complexity
Cytoplasmic Specification
Cells developmental fate determined by cytoplasmic components (proteins, mRNA's) (cell fate)
Mosaic Development
Process in early animal development that independently differentiates each part of embryo. (cell fate)
Conditional Specification
Cell is induced to a particular fate by neighboring cells; cell fate changes if moved in embryo. (cell fate)
Syncytial Specification
Molecules that influence cell fate diffuse within the the cytoplasm of a single large, multi-nucleated cell. Seen in Arthropods (cell fate)
Morphogens
A signaling molecule that controls tissue development and pattern formation by spreading through tissue to establish gradients in the embryo.
Segmentation
Serial repetition of similar body regions along the anteroposterior axis of the body. ex. worms, fish
Metamerism
2 colors match under one specific light source but look different when light changes. (only 3 phyla have: annelids, arthropods, chordata)
HOX genes
Plan and placement of body parts during embryonic development. Master genes that specify a location within the body.
Radial Cleavage
Cleavage planes are symmetrical to the polar axis, producing layers of cells on top of each other, regulative.
Regulative Development - no fixed fates initially, communicate with neighboring cells and environment
ex: humans, fish, sea urchins
Spiral Cleavage
Cleaves oblique (off-center of axis) to axis and typically produces a quartet of cells that lie in furrows between the cells; mosaic.
Mosaic Development - predetermined cell fate
ex: snails, worms
Blastula
Hallow ball of cells that forms during early animal embryonic development.
Blastocoel
Ball of cells surrounding a fluid-filled cavity
Gastrula
Hallow ball of cells (blastula) reorganizes into a structure with multiple tissue layers
Ectoderm
The outer derm layer surrounding blastocoel
Endoderm
The inner layer of blastocoel, surrounding inner cavity.
Gastrocoel
The inner body cavity of a blastocoel
Diploblasts, Triploblasts
2 layers, 3 layers of tissue
Coelom
A fluid-filled body cavity that is completely lined with tissue derived from the mesoderm and surrounds the digestive tract and internal organs. Allows for protection (shock absorber), movement (shift shape), support (hydrostatic), circulation (transporting gases, nutrients, waste products)
Pseudocoelom (type of body cavity)
A fluid filled cavity that lies between the body wall and the digestive tract. Mesoderm lines the outer edge of the blastocoel.
Acoelomate (type of body cavity)
No body cavity, the space between the digestive tract and outer body cells is completely filled with tissue. Mesoderm completely fills the blastocoel.
Coelomate (type of body cavity)
A fluid filled cavity New cavity completely lined with a middle tissue layer called mesoderm. MOST SA - has largest body cavity.
Schizocoely (way to form gut)
An embryonic development process that interally splits to form the fluid-filled body cavity. Mesodermal cells migrate to blastocoel to form the coelom.
Enterocoely (way to form gut)
Coelom pinch off from pouches of the gut that push outward into the blastocoel.
Triploblastic Bilateria
Phyla that consist of bilateral animals with three germ layers
Protostomia
Form of embryonic development where "mouth first"=blastopore becomes mouth, spiral mosaic cleavage, triploblastic, coelomates form via schizocoely
Ecdyosozoa
Molt their cuticles - shedding animals
Lophotrochozoa
Bilateral symmetry, spiral cleavage, triploblastic animals with protostomes(first opening in embryo develops into the mouth)
Most have trochophore larval stage - free-swimming stage, moving via cillia
Deuterostomia
"Mouth second" = blastopore become the anus, radial regulative cleavage, all are coelomates, vertebrate members form their coelom via schizocoely, Enterocoelous
Taxonomy
Formal system for naming and classifying species
Systematics
Broader science of comparative biology
Kingdom, Phylum, Class, Order, Family, Genus, and Species (King Phillip Came Over For Good Soup)
Taxa
Names of groups at each rank in the biological classification system to categorize organisms
Binomial Nomenclature
Linnaeus's system for naming species
Phylogeny
The evolutionary history of an organism or taxa of organisms
Characters
Studying organismal features, including: morphological, biochemical, chromosomal, and molecular features.
Homoplasty
Character similarity that misrepresents common descent (looks similar but misrepresents descent)
ex: bird and bat wings
Ancestral character state
The character state present in the common ancestor.
Evolutionary derived character state
Variant forms of the character traits that arose later within the group
Polarity of a character
The ancestral/descendant relationships among the variant forms of a character.
Outgroup
Group that is phylogenetically close but not within the group being studied.
Principle of Parsimony
Prefer the simplest hypothesis capable of explaining the observations. Simplest explanation is the correct one.
Clade
Species that share derived character states form subsets within the study group.
Synapomorphy
Derived character shared by members of a clade. Used as evidence of homology. Finding a clade then we know the organisms are all related to each other.
Nested Hierarchy
Pattern of derived states of characters within study group forms clades within clades.
Smaller groups fit completely inside larger groups.
Monophyletic
Includes the most recent common ancestor and all descendants of that ancestor.
Paraphyletic
Includes the most recent common ancestor of all members of a group and some but not all descendants of that ancestor.
Polyphyletic
Does not include the most recent common ancestor of all members of a group; has at least 2 separate evolutionary origins.
Traditional Evolutionary Taxonomy
Accounts both for common descent and adaptive evolutionary change. Taxa may be monophyletic or paraphyletic, REJECTS polyphyletic grouping.
- A distinct adaptive zone may merit its own taxon (ex. Hominidae)
Phylogenetic Systematics or Cladistics
Emphasizes common descent and cladograms. Rejects paraphyletic and polyphyletic groups. (only likes evolutionary history, common ancestors evolving)