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BIO 16 ACTIN

front 1

what does actin form

back 1

microfilaments

front 2

where is actin most likely present

back 2

cortex, below the plasma membrane

front 3

what are the plural forms of actin

back 3

lamellipodia and filopodia (F-actin)

front 4

what is the monomer of actin

back 4

G-actin

front 5

How does actin exibit polarity

back 5

has a minus and plus end - show speed of shrinkage or growth

front 6

What is the function of nucleations

back 6

allows for growth

front 7

What happens in the step of nucleation?

back 7

the lag state - 3 G-actins form a nucleus (trimer)

Slow step forms the nucleus

front 8

What happens in the elongation state?

back 8

the growth phase - actin filament - rapid growth once nucleus formed

front 9

What happens in the steady state?

back 9

the equilibrium phase - subunits being added and subtracted at equal rates

rate of association = rate of dissociation (Cc)

front 10

What is the significance of the plus and minus end of actin?

back 10

(-) end has a higher critical concentration

growth and shrinkage can occur on both ends

front 11

What is treadmilling?

back 11

growth on one end, and shrinkage on the other = no change in length

both ends are constantly dynamic

front 12

How is the ATP cap formed?

back 12

when the rate of G-actin added is greater than the rate of ATP hydrolysis

- means constant growth

front 13

what form of ATP when added?

back 13

ATP added

ADP removed

front 14

What are the three toxins for actin

back 14

Latrunculin - depolymerizes

Cytochalasin B - depolumerizes

Phalloidin - stabilizes (unable to disassemble)

front 15

What are the three toxins for microtubules

back 15

Taxol - stabilizes (unable to disassemble)

Nocodazole - depolymerizes

Colchicine - depolymerizes

front 16

What are the 2 proteins that modify G-actin?

back 16

Thymosin- inhibit addition of G-actin to bind on plus end

Profilin - favors addition of G-actin to bind to plus end

front 17

What are the 2 examples of an actin nucleator?

back 17

Arp 2/3: (branched actin) caps the minus end so only growth occurs on the plus end

Formin: (unbranched actin) see-saw back and forth - bound to plus end and adding G-proteins on the plus end

front 18

What is the function of CapZ?

back 18

prevents growth and shrinkage on the plus end (cap)

front 19

What is the function of the tropomodulin?

back 19

prevents growth and shrinkage on the minus end (cap)

front 20

what is tropomodulin?

back 20

a type of capping protein

front 21

Where can capping proteins be found?

back 21

skeletal muscle cells

front 22

what is the significance of these actin filaments being capped?

back 22

this ensures only sliding is occurring between the different actin forms (no shrinking or growing)

front 23

What is the function of cofilin?

back 23

an actin severing protein - forces filament to twist tightly until breaks - allows for broken actin to be recycled and added to the top as ATP

front 24

What are the 2 actin bundling proteins?

back 24

fimbrin - shorter protein, holds actin closer together

Alpha-actinin - longer protein, holds actin farther apart

front 25

What type of bundling protein is used for a contractile bundle?

back 25

alpha - actinin (need more space to but motor protein between them)

front 26

what type of bundling protein is used for a tight parallel bundle?

back 26

fimbrin (prevents myosin II from entering bundle)

front 27

What is the motor protein for actin?

back 27

myosin

front 28

What is the structure of the actin motor protein that creates the contractive properties?

back 28

skeletal muscle myosin II - 2 heavy chains and 4 light chains - in a coiled coil (alpha-helical domain)

- flower like ends that join their stems creating a bare zone in the middle

front 29

what is the function of active myosin?

back 29

motor fixed, filament is not

1. in rigor state (fixed) myosin head bound to actin filament

2. ATP molecule binds to myosin head (loses affinity to actin filament)

3. myosin head hydrolyzes ATP and in cocked position

4. myosin head loses inorganic phosphate and has high affinity to bind to actin filament

5. loses ADP and changes conformation - in straight position and slides the space to a G-actin)

front 30

when is the myosin II muscle contraction active?

back 30

Ca2+ released

front 31

What are the functions of the Z-discs?

back 31

there are 2 z-discs on both ends of a sarcomere.

this creates the contractile unit of a skeletal muscle cell that shortens length by sliding

front 32

What are the 2 proteins that regulate skeletal muscle contraction?

back 32

Tropomyosin - binds filament groove, covering the myosin binding site (no nerve signal bc in the way)

Troponin - binds with Ca2+ and pulls tropomyosin out of myosin binding site so contraction can occur (nerve receives signal)

front 33

What activates myosin II contraction?

back 33

phosphorylation of the light chains

(starts in a folded inactive state, then gets phosphorylated and changes confirmation to form a baby filament)

front 34

What does the barbed end and the pointed end mean in actin filament?

back 34

pointed end - plus side

barbed end - minus side

front 35

What end is the critical concentration point always higher?

back 35

minus end

front 36

what does it mean if you have a ATP cap?

back 36

that actin is actively growing

front 37

How does formin grab G-actins?

back 37

by working with profilin which is a protein that favors the addition of G-proteins on the plus end

front 38

Where do we find branched actin?

back 38

in the cortex

front 39

Where do we find unbranched actin?

back 39

in sweeping/ crawling

- stress fibers, filopodia (finger-like projection on the surface of cells), and contractile ring

front 40

what is the cortex

back 40

in side the cell near the membrane that helps the cell with structural support and shape